Proskauer, J.M. (1957), Studies on Anthocerotales V. Phytomorphology 7: 132 [tax. nov.]

Duff, R.J., Villarreal, J.C., Cargill, D.C. & Renzaglia, K.S. (2007), Progress and challenges toward developing a phylogeny and classification of the hornworts. The Bryologist 110(2): 238, 239 [secondary reference]

Renzaglia, K.S., Villarreal, J.C. & Duff, R.J. in Goffinet, B. & Shaw, A.J. (ed.) (2009), New insights into morphology, anatomy and systematics of hornworts. Bryophyte Biology Edn. 2: 142 APC [secondary reference]

Söderström, L., Hagborg, A., Konrat, M. von, Bartholomew-Began, S., Bell, D., Briscoe, L., Brown, E., Cargill, D.C., Costa, D.P., Crandall-Stotler, B.J., Cooper, E.D., Dauphin, G., Engel, J.J., Feldberg, K., Glenny, D., Gradstein, S.R., He, X., Heinrichs, J., Hentschel, J., Ilkiu-Borges, A.L., Katagiri, T., Konstantinova, N.A., Larrain, J., Long, D.G., Nebel, M., Pócs, T., Puche, F., Reiner-Drehwald, E., Renner, M.A.M., Sass-Gyarmati, A., Schafer-Verwimp, A., Segarra Moragues, J.G., Stotler, R.E., Sukkharak, P., Thiers, B.M., Uribe, J., Vana, J., Villarreal, J.C., Wigginton, M., Zhang, L. & Zhu, R.-L. (2016), World checklist of hornworts and liverworts. Phytokeys 59: 27 [secondary reference]

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