Nakai, T. (1943), Chosakuronbun Mokuroku: Ordines, Familiae, Tribi, Genera, Sectiones. Novis Edita. Appendix: 200 [tax. nov.]

Crandall-Stotler, B., Stotler, R.E. & Long, D.G. (2009), Phylogeny and classification of the Marchantiophyta. Edinburgh Journal of Botany 66(1): 189 [secondary reference]

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• CAB Dist.: WA, SA, NSW, Vic

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