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McCarthy, P.M. (2003), Catalogue of Australian Liverworts and Hornworts. Flora of Australia Supplementary Series 21: 48 APC [secondary reference]

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Crandall-Stotler, B., Stotler, R.E. & Long, D.G. (2009), Phylogeny and classification of the Marchantiophyta. Edinburgh Journal of Botany 66(1): 189 [secondary reference]

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